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Family: Bryaceae
arctic bryum moss, more...bryum moss
[Bryum alpicola Broth., moreBryum arcticum (R. Br.) Bruch & Schimp., Bryum arcticum f. aurimontanum (Kindb.) Podp., Bryum arcticum f. flavescens (J.J. Amann) Podp., Bryum arcticum f. kaurinii (H. Philib.) Podp., Bryum arcticum f. kindbergii (H. Philib.) Podp., Bryum arcticum f. oxystegium (I. Hagen) Podp., Bryum arcticum f. submicans (Kindb.) Podp., Bryum arcticum subsp. arcuatum (Limpr.) J.J. Amann, Bryum arcticum subsp. callistomum J.J. Amann, Bryum arcticum subsp. flavescens J.J. Amann, Bryum arcticum subsp. helveticum (H. Philib.) J.J. Amann, Bryum arcticum subsp. inflatum (H. Philib.) J.J. Amann, Bryum arcticum subsp. kindbergii (H. Philib.) J.J. Amann, Bryum arcticum subsp. kongsvoldense (H. Winter) J.J. Amann, Bryum arcticum subsp. lawersianum (H. Philib.) J.J. Amann, Bryum arcticum subsp. laxirete (Bryhn & Ryan) Podp., Bryum arcticum subsp. oxystegium (I. Hagen) J.J. Amann, Bryum arcticum subsp. tomentosum (Jørg.) J.J. Amann, Bryum arcticum subsp. viride (H. Philib.) J.J. Amann, Bryum arcticum subsp. watzmannii (H. Winter) J.J. Amann, Bryum arcticum var. arcuatum (Limpr.) Podp., Bryum arcticum var. callistomum (J.J. Amann) C.E.O. Jensen, Bryum arcticum var. flavescens (J.J. Amann) C.E.O. Jensen, Bryum arcticum var. helveticum (H. Philib.) Husn., Bryum arcticum var. kindbergii (H. Philib.) C.E.O. Jensen, Bryum arcticum var. oxystegium (I. Hagen) H. Winter, Bryum arcticum var. purpurascens (R. Br.) Ångstr., Bryum arcticum var. tomentosum Jørg., Bryum arcuatulum Kindb., Bryum arcuatum Limpr., Bryum aurimontanum Kindb., Bryum autumnale Limpr., Bryum brownii Bruch & Schimp., Bryum brownii subsp. serotinum (Lindb.) J.J. Amann, Bryum bruntonii Schimp., Bryum bryoides (R. Br.) Ångstr., Bryum campylocarpum Limpr., Bryum cernuum subsp. campylocarpum (Limpr.) Podp., Bryum cirrhatum subsp. alpicola (Broth.) Podp., Bryum corioideum Bryhn & Ryan, Bryum helveticum H. Philib., Bryum inclinatum subsp. hyperboreum Podp., Bryum inclinatum var. corioideum (Bryhn & Ryan) Podp., Bryum inclinatum var. liliputanum Podp., Bryum inclinatum var. parvum (Bryhn & Ryan) Podp., Bryum inclinatum var. semiovatum Podp., Bryum inflatum H. Philib., Bryum kaurinii H. Philib., Bryum kindbergii H. Philib., Bryum kindbergii subsp. helveticum (H. Philib.) Kindb., Bryum kongsvoldense H. Winter, Bryum lawersianum (R. Br.) Bruch & Schimp., Bryum laxirete Bryhn & Ryan, Bryum lindgrenii Schimp., Bryum lisae var. alpicola (Broth.) Margad., Bryum luridum R. Ruthe, Bryum micans Mitt., Bryum micans subsp. inflatum (H. Philib.) Kindb., Bryum opdalense Limpr., Bryum opdalense subsp. tromsoeense (G. Roth) J.J. Amann, Bryum opdalense var. tromsoeense (G. Roth) H. Winter, Bryum oxystegium I. Hagen, Bryum pallens subsp. lawersianum (H. Philib.) Podp., Bryum parvifolium Cardot, Bryum parvum Bryhn & Ryan, Bryum pearyanum Bryhn, Bryum pendulum var. kaurinii (H. Philib.) J.J. Amann, Bryum purpurascens (R. Br.) Bruch & Schimp., Bryum purpurascens subsp. autumnale J.J. Amann, Bryum purpurascens subsp. lindgrenii (Schimp.) J.J. Amann, Bryum purpurascens subsp. opdalense (Limpr.) Kindb., Bryum purpurascens subsp. serotinum J.J. Amann, Bryum purpurascens var. autumnale (Limpr.) C.E.O. Jensen, Bryum purpurascens var. lindgrenii (Schimp.) C.E.O. Jensen, Bryum purpurascens var. opdalense (Limpr.) C.E.O. Jensen & Perss., Bryum purpurascens var. serotinum (Lindb.) C.E.O. Jensen, Bryum purpurascens var. tromsoeense (G. Roth) C.E.O. Jensen, Bryum rufum Fergusson, Bryum rutilans subsp. pearyanum (Bryhn) Podp., Bryum rutilans subsp. subrutilans (Kindb.) Podp., Bryum serotinum Lindb., Bryum submicans Kindb., Bryum subrutilans Kindb., Bryum tromsoeense G. Roth, Bryum uliginosum subsp. campylocarpum (Limpr.) Podp., Bryum viride H. Philib., Bryum warneum subsp. viride (H. Philib.) J.J. Amann, Bryum watzmannii H. Winter, Hemisynapsium arcticum (R. Br.) Brid., Hemisynapsium arcticum var. purpurascens (R. Br.) Brid., Hemisynapsium bryoides (R. Br.) Brid., Plagiobryum purpurascens (R. Br.) N. Pedersen, Pohlia arctica R. Br., Pohlia arctica var. purpurascens (R. Br.) Schwägr., Pohlia bryoides R. Br., Pohlia purpurascens R. Br., Ptychostomum bryoides (R. Br.) J.R. Spence, Ptychostomum purpurascens (R. Br.) J.R. Spence] |
Plants: in dense or open turfs, red, pink, green, or yellow-green. Stems: 0.5–3 cm, fertile stems comose, innovations evenly foliate. Leaves: shiny yellow-green to red or pink, crowded, strongly contorted to shrunken when dry, ovate-lanceolate, flat, 0.5–3 mm, often gradually enlarged toward stem apex; base usually green, not decurrent; margins revolute proximally, limbidium strong, in 2 or 3 rows; apex acute to acuminate; costa excurrent to long-excurrent, awn smooth or denticulate; proximal laminal cells long-rectangular, 3–5:1; medial and distal cells 15–25 µm wide, 2–3:1, walls thin to moderately thick. : Specialized asexual reproduction absent. Sexual: condition polyoicous, variable, often populations either synoicous or only archegoniate. Seta: brown or red-brown, 1–3 cm, slender, straight to somewhat flexuose. Capsule: brown, shape highly variable, elongate-pyriform to clavate, symmetric to somewhat curved, 3–5 mm, mouth orange-brown to red; hypophysis slender; operculum short-conic, weakly apiculate; peristome variable, reduced; exostome teeth yellow or brown, hyaline distally, lamellae straight, sometimes with small pores near base along mid line; endostome weakly adherent to exostome, basal membrane low to 1/2 exostome height, segments with slitlike perforations, cilia absent or rudimentary. Spores: with size often variable in same capsule, 24–30(–32) µm, yellow. Phenology: Capsules mature Jul–Aug. Moist soil banks, wet soil. low to high elevations (0-4500 m). Greenland, Alta., B.C., Man., Nfld. and Labr., Que., Alaska, Calif., Mont., Nev., Oreg., Wash., Wyo., Eurasia. Ptychostomum arcticum is one of the most variable and confusing species in the genus. Much of this variation has been described as a series of very closely related species, including P. bryoides and P. purpurascens, differing in minor details of the leaves and capsule. D. T. Holyoak (2004) synonymized several of these segregates with P. arcticum, which is here tentatively accepted. The segregates that have been reported from the flora area are distinguished below based on the work of E. Nyholm (1986+, fasc. 3). Their distribution and ecology remain poorly understood, but P. purpurascens is reported from arctic North America, while P. bryoides is reported from Greenland. Capsules are needed for identification. Ptychostomum purpurascens is immediately distinguished by the distal laminal cells 22–35 µm wide, capsules elongate-pyriform and symmetric, exothecial cells transversely rectangular, in 2–4 rows, and exostome teeth with small holes present along the mid line at base. Ptychostomum arcticum and P. bryoides contrast by distal laminal cells 17–24 µm wide; capsules clavate to ovate, symmetric or curved, exothecial cells quadrate, in 2–6 rows, and exostome teeth lacking small holes. |