Geographic distribution: Hattori (1982b) summarised the known distribution as: New Guinea, Sumatra, Java, Borneo, the Philippines, and also cited the Soloman Islands as a new record at that time.
Notes: The typical phase of this variety has distinct mammillate lobuli, with the individual cells abruptly raised into tuberculate projections, particularly those cells towards the head of the lobule.However, Hattori (1982b) noted that var. minutissima was highly polymorphic and at least in New Guinea was much more common than the type variety; Hattori added that there was much variation in the projections on the surfaces of the leaf-lobules, from “densely and highly mammillose” to “weak, subconic projection”. Further investigation of ample material is required to assess if there is any correlation between the lobule surface and any other characters that have not been explored to date, e.g., those features associated with the sporophyte, as well controlled growth studies should be undertaken to assess the genetic stability of the lobule projections.