Plants: medium-sized to large, yellow-green to red bronze, strongly or weakly glossy. Stems: 2–6 cm, somewhat to weakly complanate-foliate, subpinnate to irregularly branched. Leaves: erect-spreading, ovate to lanceolate; margins toothed distally; apex long-acuminate; costa double or obscure; alar cells quadrate or rectangular, enlarged, pigmented, walls thick, region differentiated in several rows, middle lamella apparent, supra-alar cells differentiated; laminal cells elongate, flexuose, smooth. Sexual: condition autoicous or dioicous. Seta: 2–4 cm. Capsule: inclined to suberect, cylindric; exothecial cell walls not collenchymatous; operculum conic. se United States, Mexico, Central America, Eurasia, Pacific Islands.